Here the discussion is about how, as an example a mother affects a child's behavior. So one of the reasons that candidate genes can be proven causes of behavior often may be that the mother or the father may have been the source of the gene and a child with the same genetics may act differently if the source of his allele was inherited from his mother or from his father.
Lets take an example: in an A male child, his mother or his father could be the source of the A genetics. If his mother is the A (or PA) he may behave differently compared to if his father is the A or PA.
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