https://www.researchgate.net/profile/Pingyuan-Gong/publication/350658540_The_BDNF_Val66Met_modulates_the_Dark_Triad_empathic_concern_and_aggression_as_mediators/links/606d6a274585159de501420a/The-BDNF-Val66Met-modulates-the-Dark-Triad-empathic-concern-and-aggression-as-mediators.pdf
Here the authors study three traits: machiavellianism, psychopathology and narcissism. They use aggression a little differently from NPA theory. For example aggression is a part of all three of the dark triad it seems. But they do find genes as causative.
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